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Steven Austad, of the Barshop Institute for Longevity and Aging Studies, at the University of Texas Health Science Center at San Antonio, has applied the longevity quotient—which he developed—to many species, including those depicted here. The quotient is determined by dividing the greatest recorded longevity for a species by the life span that would be expected based on the species's average weight. Except for humans and Brandt's bat, the scores refer to animals in captivity.