Sometime between three million and two million years ago, perhaps on a primeval savanna in Africa, our ancestors became recognizably human. For more than a million years their australopithecine predecessors—Lucy and her kind, who walked upright like us yet still possessed the stubby legs, tree-climbing hands and small brains of their ape forebears—had thrived in and around the continent’s forests and woodlands. But their world was changing. Shifting climate favored the spread of open grasslands, and the early australopithecines gave rise to new lineages. One of these offshoots evolved long legs, toolmaking hands and an enormous brain. This was our genus, Homo, the primate that would rule the planet.